Original Singer: Peter Gabriel
Reason for Departure: Gabriel announced his departure in the middle of the prog-rock band's tour for its crowning achievement, "The Lamb Lies Down on Broadway," citing tensions within the band and strains in his personal life.
The Replacements: Phil Collins, Ray Wilson
Following Gabriel's departure, the band reportedly auditioned 400 singers. Satisfied with none of them, Genesis chose one of its own as its new frontman: drummer Phil Collins. A much more polished, pop-leaning sound followed, as did millions in album sales. When Collins left the band in 1996, the remaining members briefly recruited Stiltskin singer Ray Wilson for its final (and ill-fated) album. —Jeff Royer
(Photos: Graham Wood, Getty Images; Dave Hogan, Getty Images)
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