THEN: Nichelle Nichols as 'Nyota Uhura'
Nichelle Nichols played Lieutenant Nyota Uhura, the communications officer aboard the USS Enterprise. Uhura famously kissed William Shatner's character, Captain James T. Kirk, on a 1968 episode of the series, marking the first-ever inter-racial kiss on United States television. —XFINITY Entertainment Staff (Photo: NBC)
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